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Along the Delaware River, where limestone juts from the shore and groundwater seeps over the rock, grow little prairies. These Calcareous Riverside Outcrops and Seeps hold the same tall grasses as the Midwest's prairies — big bluestem, Indian grass, switchgrass, little bluestem — in strips far too small for bison. They are globally rare plant communities, and in Delaware Water Gap National Recreation Area they support 26 plant species that are rare in the state. The longest is near the Dingmans Ferry bridge.

Many unique native plants grow on the Calcareous Riverside Outcrops and Seeps in the Delaware Water Gap National Recreation Area

A calcareous riverside outcrop in Delaware Water Gap National Recreation Area. Credit: Perles, National Park Service.

What kept them open — and what's changing

For a long time, the river kept these prairies open itself. Every spring, huge blocks of ice riding floods swollen with snowmelt scraped off any trees and shrubs that sprouted in the rock crevices. As the climate warms, the Delaware will see shorter, milder winters with fewer freezing days — less river ice and less scouring. Without it, trees and shrubs, especially invasive ones, need to be removed every 5 to 10 years to keep the habitat open.

Invasive plants are among the biggest threats. The problem is not where they come from but how they behave: they spread fast and crowd out native plants, especially rare ones that need special habitat.

View of a transect showing exotic shrubs forming a dense shady thicket.

Invasive shrubs have turned parts of the outcrop into a dense, shady thicket. Credit: National Park Service.

View of a transect showing an open sunny prairie-like outcrop.

An open, sunny, prairie-like outcrop. Credit: National Park Service.

Two decades of monitoring

In 1998, park staff and biologists from the New Jersey Department of Environmental Protection's Office of Natural Lands Management set up 150 plots at the Dingmans Ferry outcrop. Scientists from the Park Service's Eastern Rivers and Mountains Network revisited them in 2010 and 2020.

Monitoring which plants are present and thriving in the Calcareous Riverside Outcrops and Seeps provide clues to the health of the rare community.

Monitoring which plants are present and thriving in the Calcareous Riverside Outcrops and Seeps provide clues to the health of the rare community. NPS / Callahan.

Change, 1998–2020
New invasive species15, including aggressive shrubs such as autumn olive, bush honeysuckles and multiflora rose, which turned open outcrop into shady thicket
Biggest gainerreed canarygrass, threatening to turn the rare community into a common, weedy Reed Canarygrass Floodplain Grassland spreading along mid-Atlantic rivers
Other newcomerscommon wormwood and Japanese knotweed
Native grasses and sedgesbig and little bluestem and sedges less common
Native wildflowersmore than 140 species seen, but over half, including many asters and goldenrods, declining
Native shrubsninebark, alders, dogwoods and viburnums declining

While native plant cover has remained stabled since 1998, non-native and invasive plants now cover more area than native plants in the Dingmans Ferry Calcareous

While native plant cover has remained stabled since 1998, non-native and invasive plants now cover more area than native plants in the Dingmans Ferry Calcareous Riverside Outcrops and Seeps. NPS.

Native plant cover held steady, but non-native and invasive plants now cover more ground than natives at Dingmans Ferry.

Importance of invasive plant species at the Dingmans Ferry Calcareous Riverside Outcrops and Seeps between 1998 and 2020. Importance measures the dominance of a plant species within the community.

How dominant invasive plants became at Dingmans Ferry, 1998–2020. Credit: National Park Service.

Beetles and weevils

Some invaders have been pushed back by biological control:

  • Purple loosestrife, brought to the United States from Europe in the nineteenth century, forms dense purple stands in wetlands that crowd out native plants and degrade habitat for birds, amphibians and rare plants. Between 1996 and 2007, the park released three species of loosestrife-eating beetles. Loosestrife showed the largest decline of any plant at Dingmans Ferry, and signs of Galerucella beetles feeding were everywhere — not eliminating it, but holding it back.
  • Mile-a-minute weed, a fast, barbed vine that can smother plants, appeared in the park in the early 2000s. Staff released an approved weevil to control it, and it declined at Dingmans Ferry between 2010 and 2020.

Galerucella beetles eating purple loosestrife leaves (left) and weevil insect (Rhinoncomimus latipes) feeding on mile-a-minute weed (right)

Galerucella beetles eating purple loosestrife (left) and a weevil feeding on mile-a-minute weed (right). Credit: Shreiner, National Park Service (left); National Park Service (right).

Clearing the shrubs

Without ice to do the job, park staff, the regional Invasive Plant Management Team and an invasive plant control company have been removing autumn olive, honeysuckles and multiflora rose. By 2024, nearly all invasive shrubs had been cleared from all three sites, restoring open prairie where unique native plants can thrive.

Park staff restore the Calcareous Riverside Outcrops and Seeps at Smith's Ferry by removing invasive shrubs.

Park staff remove invasive shrubs at Smith's Ferry. Credit: National Park Service.

Sources

Based on "Park Restores Unique Riverside Habitat by Managing Invasive Plants," National Park Service Eastern Rivers and Mountains Network (Delaware Water Gap National Recreation Area), a work of the United States government in the public domain. Pictures an earlier version of this page left out were restored on 2026-09-26.

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Licence: CC0 1.0 (public domain) · Adapted from www.nps.gov

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